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Messer PW, Petrov DA.Messer PW, et al.Proc Natl Acad Sci U S A. 2013 May 21;110(21):8615-20. doi: 10.1073/pnas.1220835110. Epub 2013 May 6.Proc Natl Acad Sci U S A. 2013.PMID: 23650353Free PMC article.