1 Department of Biology Education and Kumho Life Science Laboratory, Chonnam National University, Gwangju, 61186, Republic of Korea.
2 Aquatic Plant Variety Center, National Institute of Fisheries Science, Mokpo, 59002, Republic of Korea.
3 Fisheries Seed and Breeding Research Institute, National Institute of Fisheries Science, Haenam, 58746, Republic of Korea.
4 Department of Biology Education and Kumho Life Science Laboratory, Chonnam National University, Gwangju, 61186, Republic of Korea. dwchoi63@jnu.ac.kr.