1 Sanya Nanfan Research Institute of Hainan University, Sanya, China.
2 Sanya Nanfan Research Institute, Fang Zhiyuan Academician Team Innovation Center of Hainan Province, Sanya, China.
3 Key Laboratory of Tropical Horticultural Crop Quality Regulation, College of Horticulture, Hainan University, Haikou, Hainan, China.
4 School of Life Sciences and State Key Laboratory of Agrobiotechnology, The Chinese University of Hong Kong, Shatin, China.
5 College of Agriculture, Food and Natural Resources (CAFNR), Division of Plant Sciences & Technology, University of Missouri, Columbia, MO, United States.