1 Copenhagen Plant Science Center, Department of Plant & Environmental Sciences, University of Copenhagen, Frederiksberg C 1871, Denmark.
2 Max Planck Institute of Molecular Plant Physiology, Potsdam-Golm 14476, Germany.
3 Cereal Crops Research Unit, USDA-ARS, Madison, WI 53726, USA.
4 Department of Animal, Plant and Soil Sciences, School of Agriculture, Biomedicine and Environment, La Trobe University, Bundoora, VIC 3086, Australia.
5 Research School of Biology, The Australian National University, Acton, Australian Capital Territory 2600, Australia.
6 Joint International Research Laboratory of Metabolic & Developmental Sciences, State Key Laboratory of Hybrid Rice, SJTU-University of Adelaide Joint Centre for Agriculture and Health, School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, Shanghai 200240, China.