1 Department of Neuroscience, Ohio State University Wexner Medical Center, Columbus, OH, USA.
2 Faculty of Biology, Medicine and Health, The University of Manchester, Manchester, UK; Achücarro Basque Center for Neuroscience, IKERBASQUE, Basque Foundation for Science, Bilbao; Department of Neurosciences, University of the Basque Country UPV/EHU and CIBERNED, Leioa, Spain.