1 School of Ecology and Nature Conservation, Beijing Forestry University, Beijing, China.
2 CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan, China.
3 State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, China.
4 Key Laboratory of Ecology of Rare and Endangered Species and Environmental Protection (Ministry of Education), Guangxi Normal University, Guilin, Guangxi, China.
5 Guangxi Key Laboratory of Landscape Resources Conservation and Sustainable Utilization in Lijiang River Basin, Guangxi Normal University, Guilin, Guangxi, China.
6 Botany Unit, Finnish Museum of Natural History, University of Helsinki, Helsinki, Finland.
7 Museomics Research Group, Helsinki Institute of Life Science (HiLIFE), Helsinki, Finland.