1 Molecular and Cellular Neurobiotechnology, Institute for Bioengineering of Catalonia (IBEC); Department of Cell Biology, Physiology and Immunology, University of Barcelona; Network Centre of Biomedical Research of Neurodegenerative Diseases (CIBERNED), Institute of Health Carlos III, Ministry of Economy and Competitiveness, Madrid; Institute of Neuroscience, University of Barcelona, Barcelona, Spain.