Revision of the genus Atholus Thomson, 1859 (Coleoptera, Histeridae, Histerinae) from the Philippines with additional records

Ian Niel Dela Cruz; Masahiro Ôhara

doi:10.3897/zookeys.1158.100518

Revision of the genus Atholus Thomson, 1859 (Coleoptera, Histeridae, Histerinae) from the Philippines with additional records

Zookeys. 2023 Apr 12:1158:1-26. doi: 10.3897/zookeys.1158.100518. eCollection 2023.

Authors

Ian Niel Dela Cruz^{1

2}, Masahiro Ôhara¹

Affiliations

¹ Entomological Laboratory, Graduate School of Agriculture, Hokkaido University, N9, W9, Sapporo, 060-8589, Japan.
² Department of Biology, College of Mathematics and Natural Sciences, Caraga State University, Butuan City, 8600, Philippines.

Abstract

The Philippine species of the genus Atholus Thomson, 1859 are revised and re-examined based on museum as well as freshly collected specimens. Atholustorquatus (Marseul, 1854) is re-described, and SEM micrographs and illustrations of both male and female genitalia are provided. Atholusbakeri (Bickhardt, 1914) and Atholusnitidissimus Desbordes, 1925 are also re-described based on images of syntypes. Atholuspirithous (Marseul, 1873) and A.torquatus (Marseul, 1854) are new to the Philippine archipelago. Atholuscoelestis (Marseul, 1857) and A.philippinensis (Marseul, 1854) are provided with diagnostic descriptions and images. A key to the Philippine species is provided.

Keywords: Coleoptera; Histerini; SEM; genitalia; new record; taxonomy.

Ian Niel dela Cruz, Masahiro Ôhara.