1 Department of Biology, Faculty of Sciences, Lebanese University, Campus Michel Slayman Ras Maska, Tripoli, 1352, Lebanon.
2 Laboratory of Applied Biotechnology (LBA3B), Azm Center for Research in Biotechnology and its Applications, EDST, Lebanese University, Tripoli, 1300, Lebanon.
3 Laboratory of Georesources, Geosciences and Environment (L2GE), Microbiology/Tox-Ecotoxicology Team, Faculty of Sciences, Lebanese University, Campus Fanar, Jdeidet El-Matn, Beirut, 90656, Lebanon.
4 Inst Neurophysiopathol, Aix-Marseille Univ, CNRS, INP, Marseille, 13385, France.