1 State Key Laboratory of Microbial Technology, and Marine Biotechnology Research Center, Shandong University, Qingdao, 266237 China.
2 College of Bioengineering, Qilu University of Technology (Shandong Academy of Sciences), Jinan, 250353 China.
3 Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology, Qingdao, 266237 China.
4 College of Marine Life Sciences, Frontiers Science Center for Deep Ocean Multispheres and Earth System, Ocean University of China, Qingdao, 266003 China.