1 CNRS, UMR7178, 67087 Strasbourg, France. Electronic address: francois.criscuolo@iphc.cnrs.fr.
2 CNRS, UMR7178, 67087 Strasbourg, France.
3 CNRS, UMR7178, 67087 Strasbourg, France; School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Witwatersrand, South Africa.