1 Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration Biodiversity Conservation Key Laboratory of Sichuan Province, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China.
2 Zoige Peatland and Global Change Research Station, Chinese Academy of Sciences, Hongyuan 624400, China.
3 Key Laboratory of Geographical Processes and Ecological Security in Changbai Mountains (Ministry of Education), School of Geographical Sciences, Northeast Normal University, Changchun 130024, China.
4 State Key Laboratory of Black Soils Conservation and Utilization, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, China.
5 College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 101408, China.
6 State Key Laboratory of Tibetan Plateau Earth System, Resources and Environment (TPESRE), Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China.