1 Birmingham Medical School, College of Medical and Dental Sciences, University of Birmingham, Birmingham, UK.
2 School of Life Sciences, University of Warwick, Coventry, UK.
3 Institute of Immunology and Immunotherapy, College of Medical and Dental Sciences,University of Birmingham, UK; Department of Infectious Disease and Tropical Medicine, University Hospitals Birmingham NHS Foundation Trust, Birmingham, UK.
4 Department of Infectious Disease and Tropical Medicine, University Hospitals Birmingham NHS Foundation Trust, Birmingham, UK.