1 Key Laboratory of Alpine Ecology and Biogeography, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China. Electronic address: xcao@itpcas.ac.cn.
2 Beijing Key Laboratory of Resource Environment and GIS, College of Resource Environment and Tourism, Capital Normal University, Beijing 100048, China.
3 Institute of Geological Sciences, Palaeontology, Free University of Berlin, Berlin 12249, Germany.