1 Key Laboratory of Alpine Ecology and Biodiversity, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China; Department of Wood Sciences, Faculty of Agriculture and Natural Resources, Karaj Branch, Islamic Azad University, Karaj 3148635731, Iran.
2 Key Laboratory of Alpine Ecology and Biodiversity, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China; CAS Centre for Excellence in Tibetan Plateau Earth Sciences, Beijing 100101, China. Electronic address: zhuhf@itpcas.ac.cn.
3 Key Laboratory of Alpine Ecology and Biodiversity, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China.
4 Department of Forestry Forest Economics, Faculty of Natural Resources, University of Tehran, Karaj 3158777871, Iran.
5 Key Laboratory of Alpine Ecology and Biodiversity, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China; Department of Botany, Bacha Khan University Charsadda, Khyber PakhtunKhwa 24420, Pakistan.
6 Key Laboratory of Alpine Ecology and Biodiversity, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China; CAS Centre for Excellence in Tibetan Plateau Earth Sciences, Beijing 100101, China.