1 Division of Digestive Diseases, Department of Metabolism, Digestion, and Reproduction, St Mary's Hospital Campus, Imperial College London, London W2 1NY, UK. Electronic address: b.mullish@imperial.ac.uk.
2 MRC Centre for Molecular Bacteriology and Infection, Flowers Building, Imperial College London, London, UK.
3 Division of Digestive Diseases, Department of Metabolism, Digestion, and Reproduction, St Mary's Hospital Campus, Imperial College London, London W2 1NY, UK.