1 Key Laboratory of Biotechnology of Salt Tolerant Crops of Hainan Province, Hainan Key Laboratory for Sustainable Utilization of Tropical Bioresources (Provincial Ministry Building State Key Laboratory Breeding Base), College of Tropical Crops, Hainan University, Haikou, Hainan Province, China.
2 Sanya Nanfan Research Institute of Hainan University, Hainan Yazhou Bay Seed Laboratory, Sanya, Hainan Province, China.
3 Department of Cell Systems and Anatomy, UT Health San Antonio, Long School of Medicine, San Antonio, Texas, USA.