1 State Key Laboratory of Tree Genetics and Breeding, Northeast Forestry University, Harbin, 150040, China.
2 College of Life Sciences, Northeast Forestry University, Harbin, 150040, China.
3 Key Laboratory of Saline-alkali Vegetation Ecology Restoration (Northeast Forestry University), Ministry of Education, Harbin, 150040, China.
4 Biochemistry Institute, University of Science, Pyongyang, 999093, Democratic People's Republic of Korea.
5 Heilongjiang Key Laboratory of Plant Bioactive Substance Biosynthesis and Utilization, Northeast Forestry University, Harbin, 150040, China.
6 School of Forestry, Northeast Forestry University, Harbin, 150040, China.
7 Key Laboratory of Orchid Conservation and Utilization of National Forestry and Grassland Administration at College of Landscape Architecture, Fujian Agriculture and Forestry University, Fuzhou, 350002, China.
8 Orchid Research and Development Center, National Cheng Kung University, Tainan, 701, Taiwan, China.
9 Department of Life Sciences, National Cheng Kung University, Tainan, 701, Taiwan, China.
10 School of Forestry, Northeast Forestry University, Harbin, 150040, China. zhangpeng@nefu.edu.cn.
11 Key Laboratory of Orchid Conservation and Utilization of National Forestry and Grassland Administration at College of Landscape Architecture, Fujian Agriculture and Forestry University, Fuzhou, 350002, China. zjliu@fafu.edu.cn.
12 State Key Laboratory of Tree Genetics and Breeding, Northeast Forestry University, Harbin, 150040, China. shenhl-cf@nefu.edu.cn.
13 School of Forestry, Northeast Forestry University, Harbin, 150040, China. shenhl-cf@nefu.edu.cn.
14 College of Life Sciences, Northeast Forestry University, Harbin, 150040, China. lyhshen@126.com.
15 Key Laboratory of Saline-alkali Vegetation Ecology Restoration (Northeast Forestry University), Ministry of Education, Harbin, 150040, China. lyhshen@126.com.