1 Department of Biology, University of Turku, FIN-20014 Turku, Finland. Electronic address: markus.rantala@utu.fi.
2 School of Population Health, University of Auckland, 1023 Auckland, New Zealand.
3 Department of Biology, University of Turku, FIN-20014 Turku, Finland.
4 Institute of Ecology and Earth Sciences, University of Tartu, 51014 Tartu, Estonia; Department of Zoology and Animal Ecology, Faculty of Biology, University of Latvia, 1004 Rīga, Latvia; Department of Biotechnology, Daugavpils University, 5401 Daugavpils, Latvia.