1 The Key laboratory of Plant Resource, Conservation and Germplasm Innovation in Mountainous Region, Ministry of Education), Collaborative Innovation Center for Mountain Ecology and Agro-Bioengineering(CICMEAB), Guizhou University, Guiyang, 550025, Guizhou, China.
2 College of Biological Sciences, Guizhou Education University, Guiyang, 550018, Guizhou, China.
3 The Key laboratory of Plant Resource, Conservation and Germplasm Innovation in Mountainous Region, Ministry of Education), Collaborative Innovation Center for Mountain Ecology and Agro-Bioengineering(CICMEAB), Guizhou University, Guiyang, 550025, Guizhou, China. hzsxjh@126.com.
4 The Key laboratory of Plant Resource, Conservation and Germplasm Innovation in Mountainous Region, Ministry of Education), Collaborative Innovation Center for Mountain Ecology and Agro-Bioengineering(CICMEAB), Guizhou University, Guiyang, 550025, Guizhou, China. gdyulifei@163.com.