1 Department of Physiology, Anatomy and Genetics, University of Oxford, Sherrington Building, Parks Road, Oxford, OX1 3PT, UK.
2 Nuffield Department of Clinical Neurosciences, University of Oxford, John Radcliffe Hospital, Oxford, OX3 9DU, UK.
3 Nuffield Department of Clinical Laboratory Sciences, National Blood Service Oxford Centre, University of Oxford, John Radcliffe Hospital, Oxford, OX3 9BQ, UK.
4 Nuffield Department of Medicine, University of Oxford, NDM Research Building, Old Road Campus, Headington, OX3 7FZ, Oxford, UK.
5 Francis Crick Institute, London, NW1 1AT, UK.
6 Department of Physiology, Anatomy and Genetics, University of Oxford, Sherrington Building, Parks Road, Oxford, OX1 3PT, UK. peter.robbins@dpag.ox.ac.uk.