1 Graduate School of Frontier Biosciences, Osaka University, Suita, Osaka 565-0871, Japan.
2 Laboratory for Cell Polarity Regulation, RIKEN Center for Biosystems Dynamics Research (BDR), Suita, Osaka 565-0874, Japan.
3 Department of Cell Biology, Department of Physics, Universal Biology Institute (UBI), and International Research Center for Neurointelligence (WPI-IRCN), the University of Tokyo, Bunkyo-ku, Tokyo 113-0033, Japan.