1 Department of Environmental and Biological Sciences, University of Eastern Finland, Yliopistonranta 1, P.O. Box 1627, 70211, Kuopio, Finland. Jukka.Luukkonen@uef.fi.
2 Department of Environmental and Biological Sciences, University of Eastern Finland, Yliopistonranta 1, P.O. Box 1627, 70211, Kuopio, Finland.
3 STUK-Radiation and Nuclear Safety Authority, Helsinki, Finland.
4 Department of Environmental and Biological Sciences, University of Eastern Finland, Joensuu, Finland.
5 Department of Molecular Biology, Cell Biology and Biochemistry, Brown University, Providence, RI, USA.
6 Institute of Biomedicine, University of Eastern Finland, Kuopio, Finland.