1 Center for Infectious Disease Dynamics, Department of Biology, Huck Institutes of the Life Sciences, Pennsylvania State University, University Park, PA, 16802, USA. ebrandell08@gmail.com.
2 U.S. Geological Survey, Northern Rocky Mountain Science Center, Bozeman, MT, 59715, USA.
3 Department of Ecology, Evolution, and Behavior, University of Minnesota, Saint Paul, MN, 55108, USA.
4 Yellowstone Center for Resources, Wolf Project, P.O. Box 168, Yellowstone National Park, WY, 82190, USA.
5 Animal Health Diagnostic Center, College of Veterinary Medicine, Cornell University, Ithaca, NY, 14850, USA.
6 Department of Veterinary Population Medicine, University of Minnesota, Saint Paul, MN, 55108, USA.
7 Ontario Ministry of Natural Resources and Forestry, Trent University, 2140 East Bank Drive, Peterborough, ON, K9L 1Z8, Canada.
8 Grand Teton National Park, P.O. Drawer 170, Moose, WY, 83012, USA.
9 U.S. Geological Survey, Northern Prairie Wildlife Research Center, 8711 37th St. SE, Jamestown, ND, 58401, USA.
10 Denali National Park and Preserve, Central Alaska Inventory and Monitoring Network, P.O. Box 9, Denali Park, AK, 99755, USA.
11 Yukon-Charley Rivers National Preserve, Central Alaska Inventory and Monitoring Network, 4175 Geist Road, Fairbanks, AK, 99709, USA.
12 Department of Environment and Natural Resources, Government of the Northwest Territories, P.O. Box 900, Fort Smith, NT, X0E 0P0, Canada.
13 British Columbia Ministry of Forests, Lands, Natural Resource Operations and Rural Development, 2000 South Ospika Blvd., Prince George, BC, V2N 4W5, Canada.
14 Environment and Natural Resources, Government of the Northwest Territories, North Slave Region, NT, X1A 2P9, Canada.
15 Department of Wildland Resources, Utah State University, Logan, UT, 84322, USA.
16 U.S. Fish and Wildlife Service, Kenai National Wildlife Refuge, P.O. 2139, Soldotna, AK, 99669, USA.
17 Division of Wildlife Conservation, Alaska Department of Fish and Game, 802 3rd Street, Douglas, AK, 99824, USA.
18 Wildlife and Habitat Branch, British Columbia Ministry of Forests, Lands, Natural Resource Operations and Rural Development, 2080 Labieux Road, Nanaimo, BC, V9T 6J9, Canada.
19 Wildlife Biology Program, Department of Ecosystem and Conservation Sciences, Franke College of Forestry and Conservation, University of Montana, Missoula, MT, 59812, USA.
20 Division of Wildlife Conservation, Dept of Fish and Game, 1300 College Road, Fairbanks, AK, 99701, USA.
21 Ross University School of Veterinary Medicine, Basseterre, West Indies, KN-03, St. Kitts and Nevis.
22 Center for Infectious Disease Dynamics, Department of Biology, Huck Institutes of the Life Sciences, Pennsylvania State University, University Park, PA, 16802, USA.