1 Manchester Royal Eye Hospital and Manchester Academic Health Sciences Centre, Manchester University NHS Foundation Trust Manchester, Manchester, M13 9WL, UK. robert.harper@mft.nhs.uk.
2 Division of Pharmacy and Optometry, School of Health Sciences, Faculty of Biology, Medicine and Health, University of Manchester, Manchester, M13 9PL, UK. robert.harper@mft.nhs.uk.
3 Manchester Royal Eye Hospital and Manchester Academic Health Sciences Centre, Manchester University NHS Foundation Trust Manchester, Manchester, M13 9WL, UK.
4 Division of Pharmacy and Optometry, School of Health Sciences, Faculty of Biology, Medicine and Health, University of Manchester, Manchester, M13 9PL, UK.
5 Bristol Eye Hospital, University Hospitals Bristol NHS Foundation Trust, Bristol, BS1 2LX, UK.
6 Division of Evolution and Genomic Sciences, School of Health Sciences, Faculty of Biology, Medicines and Health, University of Manchester, Manchester, M13 9PL, UK.
7 Division of Optometry and Visual Sciences, City, University of London, London, EC1V 0HB, UK.