1 Centre for Tropical Water and Aquatic Ecosystem Research, James Cook University, Townsville, QLD 4811, Australia. Electronic address: adam.canning@jcu.edu.au.
2 School of Agriculture and the Environment, Massey University, Palmerston North, New Zealand.
3 Centre for Tropical Water and Aquatic Ecosystem Research, James Cook University, Townsville, QLD 4811, Australia.