1 Centre for Biomedical Sciences Education, School of Medicine, Dentistry and Biomedical Sciences, Queen's University Belfast, Belfast, Northern Ireland, UK.
2 Anatomy Department, DeBusk College of Osteopathic Medicine, Lincoln Memorial University, Harrogate, TN, USA.
3 Centre for Anatomy and Human Identification, School of Science and Engineering, University of Dundee, Dundee, UK.