1 Department of Primary Care and Public Health, Watson Building, Brighton and Sussex Medical School, Falmer BN1 9PH, UK; NIHR Global Health Research Unit on Neglected Tropical Diseases, Brighton and Sussex Medical School, Falmer, UK; Evolution, behaviour and environment, School of Life Sciences, University of Sussex, Falmer, UK. Electronic address: j.middleton@bsms.ac.uk.
2 Conservation Through Public Health, Entebbe, Uganda.
3 Health in Harmony, Portland, OR, USA.
4 Evolution, behaviour and environment, School of Life Sciences, University of Sussex, Falmer, UK.