1 Sam Noble Oklahoma Museum of Natural History and Department of Biology, University of Oklahoma, Norman, OK 73072, USA.
2 Department of Ecology and Evolutionary Biology, Cornell University, Ithaca, NY 14853, USA.
3 Key Laboratory for Conserving Wildlife with Small Populations in Yunnan, Southwest Forestry University, Kunming, 650024, China. yuanzhiyongkiz@126.com.