1 Department of Biology, University of Pennsylvania, Philadelphia, PA, USA. rschultz@sas.upenn.edu.
2 Department of Anatomy, Physiology and Cell Biology, School of Veterinary Medicine, University of California Davis, Davis, CA, USA. rschultz@sas.upenn.edu.
3 The Jackson Laboratory, Bar Harbor, ME, USA. john.eppig@jax.org.