1 School of Medicine and Menzies Institute, University of Tasmania, Hobart, Australia.
2 Priority Research Centre for Healthy Lungs and School of Medicine and Public Health, Faculty of Health and Medicine, University of Newcastle, Callaghan, Australia.
3 Hunter Medical Research Institute, New Lambton Heights, Australia.
4 Translational and Clinical Research Institute, Faculty of Medical Sciences, Newcastle University Medical School, Newcastle University, Newcastle Upon Tyne, UK.