1 Centre for Cellular and Molecular Physiology, Wellcome Centre for Human Genetics, and, Translational Gastroenterology Unit, Experimental Medicine, Nuffield Department of Medicine, University of Oxford, Oxford, UK.
2 Centre for Cellular and Molecular Physiology, Wellcome Centre for Human Genetics, and, Jenner Institute, Nuffield Department of Medicine, University of Oxford, Oxford, UK.
3 Translational Gastroenterology Unit, Experimental Medicine, Nuffield Department of Medicine, University of Oxford, Oxford, UK.