1 Aquatic Systems Research Group, Department of Ecology and Resource Management, University of Venda, Thohoyandou 0950, South Africa.
2 Aquatic Systems Research Group, Department of Ecology and Resource Management, University of Venda, Thohoyandou 0950, South Africa. Electronic address: dalutatenda@yahoo.co.uk.
3 Department of Biological Science and Biotechnology, Botswana International University of Science and Technology, Palapye, Botswana.
4 School of Biological Sciences, Queen's University Belfast, 19 Chlorine Gardens, Belfast BT9 5DL, Northern Ireland, United Kingdom of Great Britain and Northern Ireland.