1 Department of Biological Sciences, Graduate School of Science, Osaka University, 1‑1 Machikaneyama‑cho, Toyonaka, Osaka, 560-0043, Japan.
2 Department of Applied Biological Science, Faculty of Science and Technology, Tokyo University of Science, 2641 Yamazaki, Noda, Chiba, 278-8510, Japan. sakataku@rs.tus.ac.jp.