1 Department of Gene Function and Phenomics, National Institute of Genetics, Shizuoka 411-8540, Japan; Department of Genetics, The Graduate University for Advanced Studies, SOKENDAI, Shizuoka 411-8540, Japan; JST-Mirai Program, Japan Science and Technology Agency, Saitama 332-0012, Japan. Electronic address: smiyagis@ac.jp.