1 State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China; State Key Laboratory of Molecular Developmental Biology, School of Biological Sciences, University of Aberdeen, Scotland, UK; CAS Centre of Excellence in Animal Evolution and Genetics, Chinese academy of Sciences, Kunming, China. Electronic address: J.speakman@abdn.ac.uk.
2 State Key Laboratory of Molecular Developmental Biology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing, China.