1 Key Laboratory of Integrated Pest Management on Tropical Crops, Ministry of Agriculture and Rural Affairs, Environment and Plant Protection Institute, Chinese Academy of Tropical Agricultural Sciences, Haikou, 571101, China.
2 Center for Terrestrial Biodiversity of the South China Sea, College of Ecology and Environment, Hainan University, Haikou, 570228, China.
3 Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, Ontario, M5S 3B2, Canada.