1 Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, Coastal Ecosystems Research Station of the Yangtze River Estuary, Shanghai Institute of Eco-Chongming (SIEC), School of Life Sciences, Fudan University, 2005 Songhu Road, Shanghai 200438, PR China. Electronic address: zhangl1222@outlook.com.
2 Karst Dynamics Laboratory, MLR & Guangxi, Institute of Karst Geology, Chinese Academy of Geological Sciences, Guilin 541004, PR China. Electronic address: ztb@karst.ac.cn.
3 Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, Coastal Ecosystems Research Station of the Yangtze River Estuary, Shanghai Institute of Eco-Chongming (SIEC), School of Life Sciences, Fudan University, 2005 Songhu Road, Shanghai 200438, PR China. Electronic address: liux14@fudan.edu.cn.
4 Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, Coastal Ecosystems Research Station of the Yangtze River Estuary, Shanghai Institute of Eco-Chongming (SIEC), School of Life Sciences, Fudan University, 2005 Songhu Road, Shanghai 200438, PR China. Electronic address: mnie@fudan.edu.cn.
5 Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources, Chinese Academy of Sciences, 11A Datun Road, Chaoyang District, Beijing 100101, PR China. Electronic address: xuxingl@hotmail.com.
6 Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, Coastal Ecosystems Research Station of the Yangtze River Estuary, Shanghai Institute of Eco-Chongming (SIEC), School of Life Sciences, Fudan University, 2005 Songhu Road, Shanghai 200438, PR China; CAS Center for Excellence in Tibetan Plateau Earth Sciences, Chinese Academy of Sciences (CAS), Beijing, 100101, PR China. Electronic address: zhshrong@fudan.edu.cn.