1 Institute of Zoology, Zoological Society of London, Regent's Park, London NW1 4RY, UK. monika.bohm@ioz.ac.uk.
2 Institute of Zoology, Zoological Society of London, Regent's Park, London NW1 4RY, UK.
3 Department of Genetics, Evolution and Environment, Centre for Biodiversity & Environment Research, University College London, Gower Street, London, WC1E 6BT, London, UK.
4 Department of Biogeography, Trier University, D-54286 Trier, Germany.
5 IUCN Species Survival Commission (SSC), Calle La Joya, Edificio Unidad Técnica del Este, Chacao, Caracas 1060, Venezuela.
6 Provita, Calle La Joya, Edificio Unidad Técnica del Este, Chacao, Caracas 1060, Venezuela.
7 Venezuelan Institute for Scientific Investigation (IVIC), Caracas 1020-A, Venezuela.