1 WorldFish, Jalan Batu Maung, Batu Maung, 11960, Bayan Lepas, Penang, Malaysia. alex.tilley@gmail.com.
2 WorldFish, Jalan Batu Maung, Batu Maung, 11960, Bayan Lepas, Penang, Malaysia.
3 Australian Research Council Centre of Excellence for Coral Reef Studies, James Cook University, Douglas, Australia.
4 European Centre for Environment and Human Health, University of Exeter Medical School, Knowledge Spa, Royal Cornwall Hospital, Truro, Cornwall, TR1 3HD, UK.
5 Department of Biological Sciences, University of Bergen, PO Box 7803, 5020, Bergen, Norway.