1 Department of Biology and Environmental Science, University of Jyväskylä, Finland.
2 Messerli Research Institute, University of Veterinary Medicine Vienna, Medical University of Vienna, University of Vienna, Veterinaerplatz 1, A-1210 Vienna, Austria; University of Vienna, Department of Integrative Zoology, Althanstrasse 14, A-1090 Vienna, Austria.
3 Department of Biology and Environmental Science, University of Jyväskylä, Finland. Electronic address: bibiana.rojas@jyu.fi.