1 UMR "Animal, Health, Territories, Risks and Ecosystems" (ASTRE, CIRAD-INRA-Université de Montpellier), Montpellier, France.
2 Center for Animal Disease Modeling and Surveillance (CADMS), Department of Medicine and Epidemiology, School of Veterinary Medicine, University of California, Davis, Davis, CA, United States.
3 Institute for Game and Wildlife Research-IREC (UCLM-CSIC), Ciudad Real, Spain.