1 Academy of Animal Science and Veterinary Medicine, Qinghai University, Xining, Qinghai, 810016, China.
2 Yak Engineering Technology Research Center in Qinghai Province, Xining, Qinghai, 810016, China.
3 Key Laboratory of Qinghai-Tibetan Plateau Animal Genetic Resource Reservation and Utilization, Southwest Minzu University, Chengdu, Sichuan, 610000, China.
4 Institute of Animal Husbandry and Veterinary Medicine, Tibet Academy of Agriculture and Animal Husbandry Science, Lhasa, Tibet, 850009, China.
5 Sichuan Academy of Grassland Sciences, Chengdu, Sichuan, 611731, China.
6 Key Laboratory of Animal Genetics, Breeding and Reproduction of Shaanxi Province, College of Animal Science and Technology, Northwest A&F University, Yangling, Shaanxi, 712100, China.