1 National Institute of Biological Sciences, Beijing 102206, China.
2 Peking University-Tsinghua University-National Institute of Biological Sciences Joint Graduate Program, School of Life Sciences, Peking University, Beijing 100871, China.
3 Graduate School of Peking Union Medical College, Beijing 100730, China.
4 Tsinghua Institute of Multidisciplinary Biomedical Research, Tsinghua University, Beijing 100084, China.