1 Veterinary Sciences Division, Agri-Food and Biosciences Institute, Belfast, United Kingdom.
2 School of Biological Sciences, Queens University Belfast, Belfast, United Kingdom.
3 Wildlife Disease Laboratory, Michigan Department of Natural Resources, Lansing, MI, United States.
4 Faculty of Medicine and Health Sciences, Division of Molecular Biology and Human Genetics, DST-NRF Centre of Excellence for Biomedical TB Research, MRC Centre for Tuberculosis Research, Stellenbosch University, Cape Town, South Africa.