1 Laboratory for Proteolytic Neuroscience, RIKEN Center for Brain Science, 2-1 Hirosawa, Wako-City, Saitama, 351-0198, Japan. shoko.hashimoto@riken.jp.
2 Laboratory for Proteolytic Neuroscience, RIKEN Center for Brain Science, 2-1 Hirosawa, Wako-City, Saitama, 351-0198, Japan.
3 Department of Functional Brain Imaging Research, National Institute of Radiological Sciences, National Institutes for Quantum and Radiological Science and Technology, 4-9-1 Anagawa, Inage-ku, Chiba-City, Chiba, 263-8555, Japan.
4 Division of Neurology, Jichi Medical University, 3311-1 Yakushiji, Shimotsuke-City, Tochigi, 329-0498, Japan.
5 Center for Gene & Cell Therapy, The Institute of Medical Science, The University of Tokyo, Tokyo, 108-8639, Japan.
6 Laboratory for Proteolytic Neuroscience, RIKEN Center for Brain Science, 2-1 Hirosawa, Wako-City, Saitama, 351-0198, Japan. saido@brain.riken.jp.
7 Laboratory for Proteolytic Neuroscience, RIKEN Center for Brain Science, 2-1 Hirosawa, Wako-City, Saitama, 351-0198, Japan. takashi.saito.aa@riken.jp.
8 Department of Neuroscience and Pathobiology, Research Institute of Environmental Medicine, Nagoya University, Nagoya-City, Aichi, 464-8601, Japan. takashi.saito.aa@riken.jp.