1 Department of Life Sciences, Whitelands College, Roehampton University, London SW15 4JD, United Kingdom. Electronic address: julia.reiss@roehampton.ac.uk.
2 Department of Life Sciences, Whitelands College, Roehampton University, London SW15 4JD, United Kingdom.
3 School of Geography, Earth and Environmental Sciences, University of Birmingham, Birmingham B15 2TT, United Kingdom.
4 Centre for Functional Ecology, Department of Life Sciences, University of Coimbra, Calçada Martim de Freitas, 3000-456, Coimbra, Portugal.