1 Department of Biology and the Integrative Program for Biological and Genome Sciences, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America.
2 Department of Biostatistics, University of Washington, Seattle, Washington, United States of America.
3 Institute for Bioethics, Health Policy and Law, University of Louisville School of Medicine, Louisville, Kentucky, United States of America.
4 Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America.
5 Departments of Population and Quantitative Health Sciences, Institute of Computational Biology, Case Western Reserve University, Cleveland, Ohio, United States of America.
6 HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America.