1 Division of Cancer Research, Medical Research Institute, Ninewells Hospital and Medical School, The University of Dundee, Dundee, DD1 9, SY, United Kingdom.
2 Department of Surgery and Cancer, Imperial College London, Hammersmith Hospital Campus, London, W12 0NN, United Kingdom.
3 Division of Cancer Research, Medical Research Institute, Ninewells Hospital and Medical School, The University of Dundee, Dundee, DD1 9, SY, United Kingdom. d.w.meek@dundee.ac.uk.