Reply to Risk and Zhu: Mixed-effects modeling as a principled approach to heritability analysis with repeat measurements

Affiliations

• ¹ Athinoula A. Martinos Center for Biomedical Imaging, Massachusetts General Hospital, Harvard Medical School, Charlestown, MA 02129; tge1@mgh.harvard.edu.
• ² Psychiatric and Neurodevelopmental Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, MA 02114.
• ³ Stanley Center for Psychiatric Research, Broad Institute of MIT and Harvard, Cambridge, MA 02138.
• ⁴ Athinoula A. Martinos Center for Biomedical Imaging, Massachusetts General Hospital, Harvard Medical School, Charlestown, MA 02129.
• ⁵ Department of Psychology, Yale University, New Haven, CT 06520.
• ⁶ Department of Psychiatry, Massachusetts General Hospital, Harvard Medical School, Boston, MA 02114.
• ⁷ Department of Psychology, Harvard University, Cambridge, MA 02138.
• ⁸ Center for Brain Science, Harvard University, Cambridge, MA 02138.
• ⁹ School of Electrical and Computer Engineering, Cornell University, Ithaca, NY 14853.
• ¹⁰ Nancy E. and Peter C. Meinig School of Biomedical Engineering, Cornell University, Ithaca, NY 14853.